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signs off on Tuesday, after 12 seasons of crime solving, evidence examining, blood, guts and, most importantly, bones.

But at the core of the long-running Fox procedural -- the “heart of the matter,” if you will -- was a partnership between forensic anthropologist Temperance Brennan (Emily Deschanel) and FBI Special Agent Seeley Booth (David Boreanaz).

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As she eventually discovers, Booth is alive, and the similarities in the skeletal structure point her to the real victim: his younger brother, Jared (Brendan Fehr).

The elder Booth is merely off trying to be the hero again, a shot in the gut his only reward for trying in vain to save state secrets and his brother’s life.

However, the finale is a big one for Booth and Brennan, who deal with serious emotional fallout after the FBI agent fakes his own death to go undercover and his partner is not informed -- beer hats in bathtubs, anyone?

The episode -- and therefore, the season -- ends with Booth assuaging Brennan’s feelings of hurt and guilt at Zach’s betrayal in a sweet personal moment that’s a preview of things to come.

Two kids and countless hurdles later, they’re still going strong.

The show’s 10th season finale, “The Next in the Last,” was written before producers knew whether or not they’d be coming back for an 11th year -- and featured Booth and Brennan stepping away from the crime-solving life as they prepared to welcome their second child.

Everything goes according to plan until Booth realizes that Brennan’s plan to prove her father’s innocence involves implicating herself in the murder instead, something he’s unwilling to go along with, even under oath.

After 12 seasons, it’s difficult to pick a seminal line for these two, but “That’s a lot of heart, Bones” is certainly in the top five.

So in the two-part season 11 opener, When charred remains turn up at the Jeffersonian, all signs point to them being Booth’s.

Unsurprisingly, Brennan refuses to accept it, throwing herself back into the work and mentally cataloguing every injury her husband sustained throughout his traumatic childhood and heroic adult life in the hopes of finding something to refute a mountain of physical evidence.

From their first episode together, Brennan and Booth’s dynamic was electric, and while the will-they-or-won’t-they of it all took nearly seven seasons to sort out (more if you count that they don’t actually tie the knot until season ), their relationship kept fans hooked for all 245 episodes.

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